In the early 1990s, Yuuta Kobayashi is a Japanese boy who plays the game Mega Man 5, but forgets to turn off his console before going to bed. This allows Mega Man, along with his friends and foes, to exit the "video game world", arriving in Japan in the "real world". As Dr. Wily attempts to conquer the real world, Mega Man teams up with the Kobayashi family to search for Dr. Wily, eventually finding him at an amusement park after Yuuta's father takes him to Tokyo. Dr. Wily orders the robots in a giant machine called Samurai Man; though Mega Man defeats it by Charge Kicking a football, Dr. Wily escapes. Yuuta summons Eddie to replenish Mega Man's energy before leaving with him and Rush to search Tokyo for Dr. Wily. Along the way, Mega Man learns about Japanese culture and customs, including the Japanese New Year, Setsubun, Hinamatsuri, and Children's Day. Eventually, they discover that Dr. Wily built a secret base inside Mount Fuji and used the volcanic energy to create Lava Man. Yuuta's sister Akane summons Proto Man and Beat to help Mega Man; while they confront the lava robots, Mega Man breaks into Wily's lab and defeats his Robot Masters. Though Wily flees in a rocket, Proto Man diverts his rocket to go back into Yuuta's video game; after celebrating Wily's defeat, the characters return to the game.

Some time later, Mega Man and Roll are pulled out of the game world by Yuuta and Akane and taken to the Obon matsuri. There, Mega Man and Yuuta carry the mikoshi in a parade, Roll learns about yukatas and makes a promise with Akane that she will wear a kimono on Shichi-Go-San, Mega Man learns about cotton candy, Roll plays Whac-A-Mole, and the kids watch the fireworks. While they are gone, Dr. Wily escapes the game world with his updated Skulker and kidnaps Proto Man. After Dr. Light repairs the time machine, Mega Man and Roll use it to travel forward one year in time, only to discover that Wily has conquered Japan and raised Tokyo above the surface. They then return to the past, where they discover that a typhoon caused by Dr. Wily and his new Typhoon Robot is responsible for this. While aiming to save Proto Man and Tokyo, Mega Man learns about Tsukimi and its dango, as well as Sports Day.

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